After the Combiner Wars ended, Cybertron started to be rebuilt. However, an undead Starscream has been reincarnated as Trypticon, wreaking havoc around him. To combat this menace, Windblade gathers up a ragtag team of Transformers, including Optimus Prime and Megatron, to resurrect an ancient ally. And while some may be forever changed by the events, others may not emerge with their sparks intact.

"组合战争"结束后，赛博坦迎来了重建时代。然而，以"铁甲龙"形态复活的亡灵红蜘蛛正在星球上肆虐。为对抗这一威胁，风刃集结了一支由擎天柱、威震天等变形金刚组成的杂牌军，试图唤醒一位远古盟友。但这场战役的代价远超想象——有人将经历永久蜕变，而另一些战士，或许连火种都无法保全……