Following a tragedy, young Tula worries about being accepted despite her family name, while a skeptical Valya struggles with the decision to take the Sisterhood vow. Years later, Valya receives a message that confirms her suspicions.

While Tula tries to keep her secret project from being discovered by the other Sisters, Desmond goes on a warpath to root out insurgents.

As Tula contends with his true identity, Valya's maneuvering leads her into an epic confrontation with an increasingly powerful Desmond.