After detecting Felldrake in Rome, Xandra takes the Caballeros to the World Tree, where her fellow gods protect the planets that grow on its branches. Xandra is disappointed to find that the gods have grown older and have given up violence, and she and the Caballeros must encourage them to find the strength to fight again when Felldrake tries to take a planet's power for his own.

After retrieving the Spark of Life that Lord Felldrake intended to steal from Mount Rushmore, the Caballeros are assigned to guard it while Xandra and the nieces are at the Institute soirée. When they get bored, they stuff it into the cabana's bear rug, bringing it to life. As the Caballeros try to catch the bear as it seeks food, Xandra crashes the soirée, Sheldgoose deals with the Institute boardmembers who demand explanations for his absences, and Daisy shows off her new boyfriend Dapper Duck to Donald's anger.

The Caballeros are dead and trapped in the Underworld. As they try to escape, they find Clinton Coot, the one who left them the cabana in his will. He explains the history of the original Caballeros and how he spent his whole life following their exploits, then helps them find a way to return to life while newly-appointed Underworld managers Sheldgoose and Felldrake attempt to stop their escape. As the nieces attempt a seance to contact the Caballeros, Xandra goes to the Underworld herself to rescue them. With her help, the Caballeros return to life on Earth, while Felldrake and Sheldgoose are kicked out of the Underworld for being bad employees.