A grisly murder stokes class division on Snowpiercer, the Great Ark Train. Melanie Cavill, the powerful head of hospitality, deputizes a dangerous rebel to help solve the killing -- Andre Layton, the world's only surviving homicide detective.

Layton uses his new position as train detective to investigate the murder while gathering intel for the revolution on the side. Meanwhile, Melanie faces a resource crisis, with potentially drastic consequences for the entire train.

Layton and Till investigate the murders. Wilford leads Miss Audrey down a dark path.