Jamie's past provides his only hope of survival following the Battle of Culloden. A pregnant Claire attempts to adjust to life in 1940s Boston.

In prison, Jamie discovers that an old foe has become the warden - and has the power to make his life hell. Claire and Frank both put their best foot forward in marriage, but an uninvited guest shatters the illusion.

Jamie is pulled into intrigue while serving as a groomsman at Helwater. In 1968, Claire, Brianna and Roger struggle to trace Jamie's whereabouts, leaving Clarie to wonder if they will ever find him.

Brianna grapples with life-changing revelations and Claire must help her come to terms with the fact that she is her father's daughter. Roger brings news that forces Claire and Brianna to face an impossible choice.