Nolan and Irisa arrive in Defiance, where the town is soon divided over the death of Luke McCawley. But a bigger threat forces everyone together: An attack by the Volge.

Rafe & Nolan search underground St. Louis while looking for an escaped convict. Meanwhile, a violent Castithan tradition causes clashes amongst the residents of Defiance.

Following the big storm, Nolan and Tommy find an American astronaut who has been in an induced sleep for the past several decades.