Takanashi is visited at work by one his 4 sisters. Hiroomi the assistant cook attempts to blackmail the staff of Wagnaria so that he won't have to work. At the same time it becomes apparent that Satou the head cook has a thing for Yachiyo the head waitress.

Takanashi takes the day off work to take care of his sick sister. The staff attempts to function without them and deal with Mahiru's fear of men with no male server to deal with the male customers.

Takanashi spends some quality time with his four quirky sisters; the boss, the drunk, the writer and the manipulative. Each in their own way is reliant on Takanashi to keep their family in order. White Day has arrived and Takanashi gifts Mahiru a bag of hair pins. This is the first White Day gift she has ever received. Later he runs into her on the street worried that she might attack some random male while walking around he escorts her home. On the way they run into his eldest sister Kazue. Mahiru get flustered and admits to "assaulting" Takanashi on a regular basis.

The girls decided to take a trip to the hot springs which will be the first time for Mahiru. Takanashi concerned about the fate of the male guests at the hot springs goes as well. In the end the entire staff ends up going only to be so late that the hot spring is closed by the time they arrive.

Kyouko is gone for a week on company business and in her absence Yachiyo attempts to "serve" Jun, which results in his heart being crushed even more. Later Nazuna sees Takanashi and Mahiru walking home together and bribes Kyouko to let her work at the restaurant for the day to spy on Mahiru. Mahiru eventually admits that she likes Takanashi, though Nazuna misinterprets this thinking that she was confirming that Takanashi was a masochist.

Yamada tells a scary ghost story at the end of work and Mahiru is to scared to go home where she would be by herself. Takanashi invites her over to spend the night at his house, Yamada follows them there, claiming to be too scared even though she was the one who told the story in the first place.

Pupura finds out about Takanashis tall sisters and is upset with him for keeping these ideal women from her. As punishment she sets him up to go on a date with Mahiru. An experiment to see how much more man acclimated Mahiru has become concludes that if she can't punch Takanashi she'll kick him in the nuts.

Takanashi and Mahiru go on their "date". The staff of the restaurant fearing for Takanashi safety concoct an elaborate plan to have him cross dress so that Mahiru would think of him as a girl and not hit him. As the date comes to a close Takanashi comes to realize his true feelings for Mahiru and Mahiru receives the compliment of her life.

Sōta's novelist sister Izumi enters a slump when she assumes he is dating Popura and will stop paying attention to her. She decides to go on a journey where she ends up meeting Mahiru who helps cheer her up. As Sōta looks after Izumi, Nazuna helps out at Wagnaria, which annoys Aoi who believes she is outshining her. After hearing about it, Nazuna puts on an act to make Aoi feel better about herself.

Mahiru meets a boy calling himself Kirio Yamada, who she believes might be Aoi's brother. As he is able to withstand her punches, Mahiru ends up conversing well with Kirio, who starts developing feelings for her, but Sōta becomes downhearted when he sees the two getting along. He begins acting strangely the next day and ends up bashing his head against the wall, leading him to think he might've become a masochist. Later, Kozue manages to get the waitresses to dress up in animal cosplay.

Sōta becomes depressed when he discovers Nazuna has grown as tall as him and has Popura act like his little sister for the day. Mahiru becomes jealous of this and works with Hiroomi to try and get Sōta to call her cute. The next day, Kirio comes to Wagnaria to look for his sister, but finds himself coming into conflict with the men, whilst Hiroomi attempts to keep him and Aoi from meeting each other.

Aoi searches for her teddy bear, Daisy, only to find that Jun had just put it in her room. Later, Yachiyo overhears a conversation between Jun and Mitsuki and assumes the two are going out with each other. As Jun notices her weird behaviour, he takes some advice from Popura to put some pressure on himself to sort things out with her.

Popura and Aoi try to help Mahiru train to cure her androphobia by dressing her up as a guy, but with no such luck. Later, Sōta and Mahiru start exchanging text messages, which causes Izumi to worry. Sōta also runs into Kirio, who manages to deduce that Mahiru has feelings for him. Later one night, Aoi gets locked out of the restaurant, but Jun manages to sort things out for her. Later, Popura and Aoi try to help out Mahiru's progress by exposing her to the other men, only for her to nearly end up punching Hiroomi. Meanwhile, Popura becomes elated when she is finally able to reach the broken items report sheet that was normally out of her reach.

Souta is given the task of reporting his family's activities to Otouru Minegishi, Kazue's ex-husband and an employee of his mother who has an annoyingly masochistic behavior. The next day, Souta has to deal with poor eyesight after his glasses break because of Aoi punching him with Daisy. while Hiroomi tries to encourage Jun to ask Kyouko out for a drink.

While Sōta remains stressed over the other day's events, Izumi, assuming he had broken up with Popura, tries to set Sōta and Mahiru up with each other, unaware of the actual circumstances between them.

As Sōta feels conflicted over his feelings towards Mahiru, Aoi, wanting Hiroomi to give her some attention, tries to obtain information on the blossoming relationships. Later, Popura gets a toothache and takes the day off, having a dream where she becomes a magical girl. Meanwhile, Kirio ends up discovering Aoi, who is revealed to be the sister he was searching for, and is met with resistance when he tries to take her back home.

Staying over at Sōta's house, Aoi explains her backstory about being constantly forced to study by her mother, inevitably pointing out the reason she ran away was because Kirio ate her natto. As the other staff members try to convince Aoi to return home over the next few days, Popura asks Kirio to bring over his mother, Kikuno, who is revealed to have trouble talking with others clearly to the point that her children need a manual to understand her.

Jun finally works up the courage to confess to Yachiyo, but it doesn't go well and Yachiyo goes into hiding. After managing to find her, Jun gives a direct confession which she finally reciprocates. Afterwards, Kyoko seems to want to stop Yachiyo making parfaits for her, saying she shouldn't be tied down to her, but Jun accepts that Kyoko is a key part of the girl he fell in love with.

As Sōta stops showing up at work, Kyoko learns that he has been spending a lot more time crossdressing because of his mother. Worried she is the cause of her problem, Mahiru confronts Sōta and tells him how she thinks of her, but Shizuka catches him and forces him to quit working at Wagnaria. Learning from Kyoko that Shizuka is having Sōta crossdress until he learns to be honest about his feelings, Mahiru heads off to talk with Shizuka, having to confront each of the Takanashi sisters along the way. As Mahiru confronts Shizuka with help from Nazuna, Popura shows up and convinces Shizuka to let Sōta keep working. Afterwards, Shizuka returns to her work while Sōta and Mahiru finally confess to each other.