It is summertime for the girls; that means one thing: comfortable summer uniforms! Tsukasa and Kagami are fraternal twins, but Konata shows them the similar and different characteristics they share. Konata, Kagami, and Tsukasa each try to put their hair into a ponytail, just to see who looks best. Unfortunately, Kagami is the only one who cannot pull it off.

The group, plus Cousin Yui and Nanako, go to the beach for some summer fun. After Yui races down the road, and Nanako gets really lost, they finally make it and spend a night in a Japanese inn. After a day at the beach, and Nanako becoming too drunk to drive, they go back to the inn for a shower and to spend another night. After arriving home, the twins have to clean their bedrooms, Konata discusses cleaning tactics with Kagami, and Miyuki is told she has to go see the dentist, or her mom is bringing the dentist to their house.

The gang comes back to school after the summer break and Konata is in a good mood because she has become hooked on Marimite. Miyuki is forced to go to school without her glasses because she broke them. Tsukasa recieves her first cell phone, but its life is very short after Tsukasa leaves it in the washing machine. On Lucky Channel, Shiraishi follows up on a fan's request for a certain hairstyle.

Konata persuades Kagami and Tsukasa to come along with her to a festival of sorts: a convention called ComiKet. There, Konata tells the twins to follow her plan of buying new manga, Kagami learns how intense some manga can be, and Tsukasa gets swept into the crowds and ends up lost.

Later that day, the girls go to another festival, with this one celebrating New Year. The girls discuss why Kagami is in a different class than everyone else, Konata draws a bad fortune, and the origin of Kagami's name is revealed.

Konata pesters her father for money, and apparently he likes when she does that. Konata wishes her online friends a happy new year, including her teacher Kuroi. Konata starts to believe that the reason why her teacher doesn't have a boyfriend is because she's always online playing games.

On Lucky Channel, it was previously announced that Akira would appear in the actual show starting with this episode, but she was unable to because she got sick.

Konata thinks about what animal each of the girls, including herself, would be. She sees herself as a fox, Tsukasa as a dog, Yutaka as a squirrel, and Kagami as a bunny.

Konata and Miyuki discuss things that can improve one's eyesight.

Konata complains about how the recent elections have been screwing up the TV schedule.

Tsukasa tries reading newspapers, but it's not long until she finds them hard to read.

Yutaka and Iwasaki's class plays dodgeball during P.E. Later, they hang out with two of their classmates, Hiyori Tamura and Patricia Martin.

On Lucky Channel, Akira and Shiraishi introduce the newest characters of the Lucky Star cast. However, Akira is mad at the fact that she still hasn't appeared in the actual show yet.

It's almost time for the school's culture festival. While the girls are preparing for their classes' themes for the festival, Patricia proposes that she and the girls in her class should do a cheerleading act for the opening ceremony of the festival. Patricia "persuades" Konata into performing the routine with her, who in turn gets Kagami, Tsukasa, Miyuki, Misao, and Ayano to do it as well.

Akira Kogami finally makes an appearance on the actual show, although she and Shiraishi are still on bad terms.

Before the opening ceremony, the girls practice the cheerleading act one last time to the song, "Motteke! Sailor Fuku!"