Lifeline, a Brussels-based resistance organisation smuggling downed RAF fliers back to Britain, is endangered when Luftwaffe Major Erwin Brandt blows one of their safehouses. The Lifeline leaders, Lisa Colbert and café proprietor Albert Foiret, are suspicious when London sends Flight Lieutenant John Curtis, who had previously gone down the line with Lisa, to help co-ordinate their activities. Brandt, meanwhile, also finds assistance from Gestapo Sturnbannfuhrer Ludwig Kessler, transferred from Berlin to crack down on the evasion lines.

Lifeline’s latest intake of evaders have been forced to leave a wounded colleague behind, hidden in a barn, but Lisa and Albert are unable to make a move due to heavy German activity in that area. The wounded evader, Sergeant Howson, is befriended and helped by a young boy, Jean-Paul Dornes, but Howson needs medical attention and Jean-Paul has to contact the resistance. Unfortunately, Jean-Paul’s mother Anna, who has raised him alone since her husband died in the fighting and is now involved with Emil Schnorr, a corporal from the local barracks, is sympathetic to the Germans.

A British bomber crash lands somewhere outside Brussels, its entire crew is apparently dead. Kessler takes this as an opportunity to infiltrate Lifeline and assigns Ernst Stoller, an officer who was born and brought up in England, to substitute for one of the dead men. Meanwhile, Monique is becoming increasingly disillusioned with her life at the Candide and relationship with Albert, and finds herself letting her guard down when she becomes involved with Stoller as part of her Lifeline work.

Lifeline once again must tread carefully when one of their safehouses is blown. Meanwhile, Van Reijn, a minister in the puppet Belgian government, wants to defect to England and contacts Curtis. Curtis tries to persuade Lisa to let Van Reijn be sent down the line, citing the propaganda value of such a defection. However, matters become complicated, as Lifeline has once more been secretly infiltrated by a German officer.

Following up a lead in their continuing investigations to crack the evasion lines, Kessler and Brandt examine the details of a recent murder case in France with Curtis, in his cover as a travelling salesman, as their prime suspect. Believing Curtis to be part of the resistance, Brandt and Kessler visit the Candide and take him away for questioning. With their efforts to help evaders becoming more and more dangerous and difficult, Albert and Lisa fear the worst, as the Germans strike right at the heart of Lifeline.