Reed and Joel bury the hatchet to face a new tornado season, but armor offers little protection when the roof blows off the new Dominator. Sean mixes up his crew and unleashes tension in the path of a west Texas twister.

The deadliest tornado of the season tears a hundred mile path across Mississippi, destroying 600 homes and the town of Yazoo City. For Reed, Joel and Chris, an epic chase turns into a race to save lives, and changes their own lives forever.

After the worst tornado in his life, Reed pursues a new force of nature in TV weather woman Ginger Zee. But chasing more than weather leaves teammates out in the cold. As twisters rip through the Ozarks, one long-time chaser calls it quits.

Sean finally gives the reigns to his meteorologist-navigator Matt Hughes, who leads the team to their best tornado intercept ever. Then triumph turns to tragedy as Storm Chasers loses one of its own.

South Dakota spins up the twister of the season, and the non-stop tornado action leaves no room for drama.