人类与机器共存的世界——完全被管制统治的“不需要感情的乐园”穹顶城市Romdo。在那个乐园都市里发生了谜一般的杀人事件，同时出现了被Cogito病毒感染的出现了自我思想的Autoreiv,负责追踪踪事件的市民情报局的年轻的职业女性莉露.梅亚在偶然中发现了有别于人类与机器人的怪物存在的秘密,，随后在她的居所的墙壁上出现了神秘的留言AWAKENING,并受到了怪物的攻击。

没有人相信莉露.梅亚所见到的，该事件被定性为STALKER(在日本称为病态的跟踪者)尾随袭击并制造了爆炸的时间，同时莉露的助手Autoreiv伊基关于此事件的资料也被消除，使得莉露确信了城市中发生的神秘事件与都市统治者之间定有关联，终于在一系列的调查下发现了PROXY的存在....与此同时，在商场中，移民青年文森特.罗被神秘的PROXY追逐着并且造成了大量市民的死伤。

因为卷入了神秘事件，无端被PROXY追杀，又被市民监管局通缉的文森特·罗自知当良民无望，偶然巧遇并是在Autoreiv皮诺的指引下，沿着通风口的回廊来到了Romdo出口处，虽被随后赶来的莉露竭力阻止，但在走投无路之下绝望的跃出了ROMDO。

由于一直生活在Romdo内，莉露在没有防护衣的保护下，感染了都市外的病菌，在医生的德莫西协助下而送回了理想都市Romdo。另外一方面，通缉犯LAW坐上了兔子(船)而往自己的故乡莫斯科前进。

Re-l recovers from her illness. Daedalus shows her the disturbing mechanisms responsible for the Dome’s continued survival. Vincent makes his way home, slowly losing his companions along the way, ending up alone with Pino. A group of infected AutoReivs is unleashed on Re-l, Iggy, and Daedalus. The episode ends with Daedalus filling out Re-l’s death certificate.

警备局长鲁尔（rule）和医生迪达罗斯·梦想（dream）的对峙。充满了隐喻和代号的一集，每个人都在追寻自己存在的意义。rule自以为代表正义，可以控制摆布dream，可事实真的如此么。。。徘徊于虚拟之乡的莉露（real），拿出所有的觉悟，寻找这世界背后的规则（law）。

An infected Iggy becomes enraged and kidnaps Re-l, setting out to kill Vincent. Re-l is again confronted by the homeless AutoReiv.

Romdo is on high alert as they search for the fugitive Raul, who plans to activate a deadly weapon. Meanwhile, the trio discover strange humanoids living inside a toxic cave system.

The trio reach the devastated Mosko, Vincent’s home city, and find that the truths lead back to Romdo.

Vincent finds himself trapped within Re-l’s consciousness, and observes powerlessly as events unfold.

A Cogito outbreak spreads through Romdo and the city falls into chaos. Vincent incites the people to revolt, pushing them to hunt down and kill all AutoReivs.

As the city continues to crumble, Vincent and Re-l encounter Proxy One. Raul is overcome with regret and despair, and sets out to find Pino.