The General will reward the one who stops Yankee Doodle Pigeon with a 30-day furlough with all expenses paid.

Fully aware of the gravity of the situation, Dick Dastardly and his proud men try to eliminate the pigeon with a feather seeking homing missile.

The General sends an efficiency expert in order to shape up the Vulture Squadron. Having analysed the results of an attitude test, he recommends Muttley to take command of the squadron.

Having messed up another mission a safe landing in a barn would be all they'd need. The country woman Hilda thinks her Prince Charming Zilly has fallen out of the sky. She won't let him go.

Yankee Doodle Pigeon and the members of the Vulture Squadron, Dick Dastardly, Muttley, Klunk and Zilly, are introduced shortly. We witness their first attempts to stop that pigeon.

Cleverly disguised as pigeons our militant men once again try to catch Yankee Doodle Pigeon. Not knowing they'd be better off as a cuckoo.

High in the sky our gallant courier Yankee Doodle Pigeon now sweeps swiftly trough the canons of the Swiss Alps. Of course, followed by Dastardly and his men who are having their planes rocked.

This time around the group try to get Yankee Doodle by shooting arrows at him. They all miss Yankee Doodle and Dastardly suggest they they need glasses. So off they go to take an eye test. They all fail and get glasses. However the glasses make their sight worse. Everything Muttley sees is farther than it really is. Klunk's vision is multiplied and he sees more than what is really there. Zilly sees everything upside down. Then The General gives Dastardly glasses and all he sees is magnified.

The General sends over a movie director to film the crew and see what they are doing wrong. The crew goes about their normal business and the director loves the footage he is getting. The crew destroys the film and the director is very upset.

Dastardly offers to give The General Proof that the squadron is doing its job and not just collecting flight pay. Muttley is put in charge of taking pictures.

The General is coming to the Vulture Squadron's hangar to inspect Yankee Doodle Pigeon having been caught. Since our inventive idiots appear to be still unable to stop that pigeon before the General arrives, they hire an actor pigeon to make the General believe it's the real Yankee Doodle Pigeon.

During 'Operation Catapault,' Dastardly falls out of the plane and lands on his head. As a result of this, he loses his memory. Klunk takes command of the squadron, but their operations are no more successful than before. Eventually, Dastardly, Zilly and Klunk all fall on their heads from a plane. Dastardly's memory is restored by this, but now Klunk and Zilly don't seem to know who he is or who they are...

The complete collection of Muttley's medals appear to be stolen. Muttley insists on Dick Dastardly helping him to get back the medals. Until then, he refuses to do his duty.

Dick Dastardly lets successively Klunk, Zilly and Muttley come up with an idea to get Yankee Doodle Pigeon. Muttley develops a bird dog sensing device that's certain stop the pigeon.

Having crocked another plane, the Vulture Squadron drops into Sam Swami's tent. He who can look into the future, predicts a better future for our fearless flyers, but a worse one for Yankee Doodle Pigeon.

The Vulture Squadron is transferred to a place where there's positively no pigeon except Yankee Doodle Pigeon.

Klunk develops a plane with a spare plane that will save our fearless flyers in case of disaster.